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rom time to time, attacks on neo-Darwinism are mounted, usually by persons who either view evolutionary theory as antireligious or who misunderstand Darwin s theory. One attack, entitled Darwin s Mistake, by Tom Bethell, was published in Harper s magazine in 1976. Bethell began by pointing out that Darwinian theory is a tautology rather than a predictive theory. (The term tautology means a statement that is true by definition.) That is, evolution is the survival of the ttest. But who are the ttest Obviously, the individuals who survive. Thus, without an independent criterion for tness other than survival, we are left with the statement that evolution is the survival of the survivors. This, indeed, is a tautology. But it is possible to assign independent criteria for fitness. Darwin wrote extensively about arti cial selection in pigeons, in which the breeders choice was the criterion for tness. (Many novel breeds of pigeon have been created this way.)

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Plant and animal breeders have practiced arti cial selection extensively. Here, survival is not the criterion for tness; productivity is. It is more dif cult to establish a priori independent criteria of fitness in nature. Often, uncontrolled or unseen vagaries have major impacts on the course of events. Surely the temperature became colder before the mammoths became woolly. Is it then reasonable to predict that elephants would get woolly if the climate became colder in Africa today The answer is no, for several reasons. First, the elephants might adapt to colder weather in any of a large number of different ways they could get fatter, they could migrate, and so on. To some extent,

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Equation 1646 includes a very important assumption: that the energy is equal to the co-energy If you make reference to Figure 168, you will realize that in general this is not true Energy and co-energy are equal only if the -i relationship is linear Thus, the useful result of equation 1646, stating that the magnetic force acting on the moving iron is proportional to the rate of change of stored energy with displacement, applies only for linear magnetic structures Thus, in order to determine the forces present in a magnetic structure, it will be necessary to compute the energy stored in the magnetic eld To simplify the analysis, it will be assumed hereafter that the structures analyzed are magnetically linear This is, of course, only an approximation, in that it neglects a number of practical aspects of electromechanical systems (for example, the nonlinear -i curves described earlier, and the core losses typical of magnetic materials), but it permits relatively simple analysis of many useful magnetic structures Thus, although the analysis method presented in this section is only approximate, it will serve the purpose of providing a feeling for the direction and the magnitude of the forces and currents present in electromechanical devices On the basis of a linear approximation, it can be shown that the stored energy in a magnetic structure is given by the expression F Wm = (1647) 2 and since the ux and the mmf are related by the expression Ni F = = (1648) R R the stored energy can be related to the reluctance of the structure according to 2 R(x) Wm = (1649) 2 where the reluctance has been explicitly shown to be a function of displacement, as is the case in a moving-iron transducer Finally, then, we shall use the following approximate expression to compute the magnetic force acting on the moving iron: dWm 2 dR(x) = dx 2 dx.

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adaptation depends not only on the changing environment, but also on the reserve variation within the gene pool of the species. Second, the elephants could become extinct; they might not be able to adapt at all. And third, if the climatic changes were not severe, the elephants might not change at all. Predicting the exact course of evolution is nearly impossible.To provide independent criteria for tness in nature is, therefore, very dif cult. Some modern evolutionary biologists, although not doubting neoDarwinism, do worry to some extent about the dif culties in testing modern evolutionary theory. However, lower-level experiments are done to test various aspects of evolution in speci c systems. For example, in 1993, B. Grant and P. Grant hypothesized that changes in bill size would occur in the nch Geospiza fortis (see g. 21.6) because of changing food size due to changing weather on the Gal pagos Islands. Their proposal seems to be correct. In addition, the support for Darwinism (the fossil

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