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dominant allele. Similarly, the (1 s1, 1 s2, 1) model is eliminated for the same reason (allele A2 is acting like the dominant allele and A1 like the recessive allele). We now describe the outcome of each of the models in the table. In both models 1 and 3 (table 20.4), selection is against genotypes containing the A2 allele. Model 1, which we just derived in detail, is the model for a deleterious recessive allele. Almost any enzyme defect in a metabolic pathway ts this model, such as PKU, alkaptonuria, Tay-Sachs disease, and so on. In model 3, however, natural selection can detect the heterozygote, as is the case with deleterious alleles that are not completely recessive. An example would be the hemoglobin anomaly called thalassemia, a disorder common in some European and Asian populations, that produces a severe anemia in homozygotes and a milder anemia in heterozygotes. It should be clear that selection can more quickly eliminate a partially recessive allele than a completely recessive allele because the allele can no longer hide in the heterozygote. Dominant or semidominant alleles (model 3) are usually more quickly removed from a population because they are completely open to selection. It takes an in nite number of generations to remove a recessive lethal allele, but only one generation for natural selection to remove a completely dominant lethal allele (see model 3, where s1 s2 1). Examples of dominant deleterious traits in

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A simple magnetic structure, very similar to those examined in the previous examples, nds very common application in the so-called variable-reluctance position sensor, which, in turn, nds widespread application in a variety of con gurations for the measurement of linear and angular velocity Figure 1623 depicts one particular con guration that is used in many applications In this structure, a permanent magnet with a coil of wire wound around it forms the sensor; a steel disk (typically connected to a rotating shaft) has a number of tabs that pass between the pole pieces of the sensor The area of the tab is assumed equal to the area of the cross section of the pole pieces and is equal to a 2 The reason for the name variable-reluctance sensor is that the reluctance of the magnetic structure is variable, depending on whether or not a ferromagnetic tab lies between the pole pieces of the magnet

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people are Huntington disease, facioscapular muscular dystrophy, and chondrodystrophy. Model 2 is interesting because selection against the heterozygote leads to an unstable equilibrium at q 0.5. If one heterozygote is removed by selection, one each of the two alleles is eliminated. However, if p and q are not equal (and thus not equal to 0.5), then one A1 allele is not the same proportion of the A1 alleles as one A2 allele is of all the A2 alleles. In other words, in a population of fty individuals with q 0.1 and p 0.9, one A2 allele is 10% (1/10) of the A2 alleles, whereas one A1 allele is only 1.1% (1/90) of the A1 alleles. Removing one each of the two alleles causes a decrease in q. Therefore, a population following model 2 is at equilibrium at p q 0.5. However, this is an unstable equilibrium. Any perturbation that changes the allelic frequencies causes the rarer allele to be selected against and eventually removed from the population. An example is the maternal-fetal incompatibility at the Rh locus in human beings. The disease erythroblastosis occurs only in heterozygous fetuses (Rh Rh ) in Rh-negative (Rh Rh ) mothers. Heterozygotes are, therefore, selected against. In model 4, selection is against homozygotes. This model is called the heterozygote advantage, and we will derive its equilibrium condition because the results are important to evolutionary theory (table 20.5). At equilibrium q pq(s1 p s2q) W

The hybrid journal bearing is superior to both axial groove and circumferential groove hydrodynamic bearings when a dynamic loading is to be applied in widely varying radial directions The disadvantage of hybrid bearings is the same as for hydrostatic bearings It is normally necessary to provide auxiliary hydraulic equipments, effective filtration and flow control restrictors However, it is possible that a lower system pressure will suffice in view of the high overload capability

The principle of operation of the sensor is that an electromotive force, eS , is induced across the coil by the change in magnetic ux caused by the passage of the tab between the pole pieces when the disk is in motion As the tab enters the volume between the pole pieces, the ux will increase, because of the lower reluctance of the con guration, until it reaches a maximum when the tab is centered between the poles of the magnet Figure 1624 depicts the approximate shape of the resulting voltage, which, according to Faraday s law, is given by eS = d dt

(20.23)

The rate of change of ux is dictated by the geometry of the tab and of the pole pieces, and by the speed of rotation of the disk It is important to note that, since the ux is changing only if the disk is rotating, this sensor cannot detect the static position of the disk

For this expression to be zero, either p 0, q 0, or (s1 p s2q) 0

Magnet Tab eS (V)

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