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Unfolded, it has a surface extent of about 4000 cm2 about the size of a full sheet of newsprint (right and left pages) Contained in the cortex are many billions of neurons (estimated at 10 to 30 billion) and ve times this number of supporting glial cells The intercellular synaptic connections number in the trillions Since nerve cells look alike and presumably function alike, the remarkable diversity in human intelligence, store of knowledge, and behavior must depend on the potential for in nite variations in neuronal interconnectivity Most of the human cerebral cortex is phylogenetically recent, hence the term neocortex It has also been referred to as isocortex (Vogt) because of its uniform embryogenesis and morphology These latter features distinguish the neocortex from the older and less uniform allocortex ( other cortex ), which comprises mainly the hippocampus and olfactory cortex.

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Concerning the detailed histology of the neocortex, six layers (laminae) can be distinguished; from the pial surface to the underlying white matter they are as follows: the molecular (or plexiform), external granular, external pyramidal, internal granular, ganglionic (or internal pyramidal), and multiform (or fusiform) layers, as illustrated in Fig 22-3 Two cell types relatively large pyramidal cells and smaller, more numerous rounded (granular) cells predominate, and variations in the lamination of the neocortex are largely determined by variations in the size and density of these neuronal types Many variations in lamination have been described by cortical mapmakers, but two main types of neocortex are recognized: (1) the homotypical cortex, in which the six-layered arrangement is readily discerned, and (2) the heterotypical cortex, in which the layers are less distinct.

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The association cortex the large areas (75 percent of the surface) that are not obviously committed to primary motor or sensory functions is generally of this latter type Homotypical areas are characterized by either granular or agranular nerve cells The precentral cortex (Brodmann s areas 4 and 6, mainly motor region) is dominated by pyramidal rather than granular cells, especially in layer V (hence the term agranular) Agranular cortex is distinguished by a high density of large pyramidal neurons In contrast, primary sensory cortex postcentral gyrus (areas 3, 1, 2), banks of the calcarine sulcus (area 17), and the transverse gyri of Heschl (areas 41 and 42) where layers II and IV are strongly developed for the receipt of afferent impulses, has been termed granular cortex because of the marked predominance of granular cells Granular cortex has a preponderance of small neurons (Fig 22-4)..

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